Now observe what very extraordinary phenomena are presented here.You have an accidental variation arising from what you may call a monstrosity; you have that monstrosity tendency or variation diluted in the first instance by an admixture with a female of normal construction, and you would naturally expect that, in the results of such an union, the monstrosity, if repeated, would be in equal proportion with the normal type; that is to say, that the children would be half and half, some taking the peculiarity of the father, and the others being of the purely normal type of the mother; but you see we have a great preponderance of the abnormal type.Well, this comes to be mixed once more with the pure, the normal type, and the abnormal is again produced in large proportion, notwithstanding the second dilution.Now what would have happened if these abnormal types had intermarried with each other; that is to say, suppose the two boys of Salvator had taken it into their heads to marry their first cousins, the two first girls of George, their uncle? You will remember that these are all of the abnormal type of their grandfather.The result would probably have been, that their offspring would have been in every case a further development of that abnormal type.You see it is only in the fourth, in the person of Marie, that the tendency, when it appears but slightly in the second generation, is washed out in the third, while the progeny of Andre, who escaped in the first instance, escape altogether.
We have in this case a good example of nature's tendency to the perpetuation of a variation.Here it is certainly a variation which carried with it no use or benefit; and yet you see the tendency to perpetuation may be so strong, that, notwithstanding a great admixture of pure blood, the variety continues itself up to the third generation, which is largely marked with it.In this case, as I have said, there was no means of the second generation intermarrying with any but five-fingered persons, and the question naturally suggests itself, What would have been the result of such marriage? Reaumur narrates this case only as far as the third generation.Certainly it would have been an exceedingly curious thing if we could have traced this matter any further; had the cousins intermarried, a six-fingered variety of the human race might have been set up.
To show you that this supposition is by no means an unreasonable one, let me now point out what took place in the case of Seth Wright's sheep, where it happened to be a matter of moment to him to obtain a breed or raise a flock of sheep like that accidental variety that Ihave described--and I will tell you why.In that part of Massachusetts where Seth Wright was living, the fields were separated by fences, and the sheep, which were very active and robust, would roam abroad, and without much difficulty jump over these fences into other people's farms.As a matter of course, this exuberant activity on the part of the sheep constantly gave rise to all sorts of quarrels, bickerings, and contentions among the farmers of the neighbourhood; so it occurred to Seth Wright, who was, like his successors, more or less 'cute, that if he could get a stock of sheep like those with the bandy legs, they would not be able to jump over the fences so readily, and he acted upon that idea.He killed his old ram, and as soon as the young one arrived at maturity, he bred altogether from it.The result was even more striking than in the human experiment which I mentioned just now.
Colonel Humphreys testifies that it always happened that the offspring were either pure Ancons or pure ordinary sheep; that in no case was there any mixing of the Ancons with the others.In consequence of this, in the course of a very few years, the farmer was able to get a very considerable flock of this variety, and a large number of them were spread throughout Massachusetts.Most unfortunately, however--Isuppose it was because they were so common--nobody took enough notice of them to preserve their skeletons; and although Colonel Humphreys states that he sent a skeleton to the President of the Royal Society at the same time that he forwarded his paper, I am afraid that the variety has entirely disappeared; for a short time after these sheep had become prevalent in that district, the Merino sheep were introduced; and as their wool was much more valuable, and as they were a quiet race of sheep, and showed no tendency to trespass or jump over fences, the Otter breed of sheep, the wool of which was inferior to that of the Merino, was gradually allowed to die out.
You see that these facts illustrate perfectly well what may be done if you take care to breed from stocks that are similar to each other.
After having got a variation, if, by crossing a variation with the original stock, you multiply that variation, and then take care to keep that variation distinct from the original stock, and make them breed together,--then you may almost certainly produce a race whose tendency to continue the variation is exceedingly strong.
This is what is called "selection"; and it is by exactly the same process as that by which Seth Wright bred his Ancon sheep, that our breeds of cattle, dogs, and fowls, are obtained.There are some possibilities of exception, but still, speaking broadly, I may say that this is the way in which all our varied races of domestic animals have arisen; and you must understand that it is not one peculiarity or one characteristic alone in which animals may vary.There is not a single peculiarity or characteristic of any kind, bodily or mental, in which offspring may not vary to a certain extent from the parent and other animals.